The Protanisoptera, another ancestral group which lacks certain wing vein characters found in modern Odonata, lived from the Early to Late Permian age until the end Permian event, and are known from fossil wings from current day United States, Russia, and Australia, suggesting they might have been cosmopolitan in distribution. Dragonfly Stay Wild Moon Child shirt The forerunners of modern Odonata are included in a clade called the Panodonata, which include the basal Zygoptera (damselflies) and the Anisoptera (true dragonflies). Today there are some 3000 species extant around the world.The relationships of anisopteran families are not fully resolved as of 2013, but all the families are monophyletic except the Corduliidae.
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The Gomphidae are a sister taxon to all other Anisoptera, the Austropetaliidae are sister to the Aeshnoidea, and the Chlorogomphidae are sister to a clade that includes the Synthemistidae and Libellulidae.
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